Swifty, an arctic fox, dreams of becoming one of the famous Top Dogs, a team of sled dogs that delivers packages for the Artic Blast Delivery Service. In the Arctic everything comes from somewhere else and the delivery service is the hub of the community. Which makes the Top Dogs the heroes of the Arctic.
In an attempt to prove his worth, Swifty secretly puts himself in command of a sled that has to deliver a mysterious package to a secret and icy fortress, where he finds himself face to face with Dr. Walrus (John Cleese), a fat walrus genius from the evil that walks on robotic legs, and commands a faithful army of bizarre trained puffins. Swifty stumbles on Dr. Walrus’ evil plan, which wants to create a tunnel to the center of the Earth to melt the polar ice caps, flooding the lava planet and becoming the king of the world.
Now Swifty has to resort to the help of his friends: PB (Alec Baldwin), an introverted polar bear, Lemmy (James Franco), a distracted albatross and rock and roll, Sal (Omar Sy) and Weez (Heidi Klum), two otters conspiracy theorists, and Jade, a red fox as hard as steel, so Swifty has a secret crush. This untidy team will have to unite and overcome their differences if it wants to prevent Dr. Walrus and his army of puffins from destroying the world. Welcome to the Great Cold North!